Borkum is the largest East Frisian Island (32 square km) in the Lower Saxony Waddensea National Park (Germany). It represents the eastern end of the German Waddensea Islands. Borkum consists of two parts which were grown together in the last centuries.
Dune succession reached the stage of old grey and brown dunes with a maximum height of more than 18 m. Borkum has the greatest variety of biotopes of all East Frisian Islands.
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Fig.1: Borkum, landscape pattern and special sites (Abbreviations: see Table 1; numbers: 1 = primary dune, 2 = white dune, 3 = Hippophae-Sambucus-zone, 4 = grassland on older dunes, 5 = dry heathland, 7= dry Salix-repens-zone, 8 = moist Salix-repens-zone, 9 = cultivated grassland, 11 = forest, 12 = transition zone, 14 = upper salt marsh, 15 = lower salt marsh, 16 = built-up area, 17 = larger bodies of water
References:
Rose, A. (2000): Die Kurzflügelkäfer der Sammlung F. und R. STRUVE von der Nordseeinsel Borkum (Coleoptera: Staphylinidae). - Ent. Bl. 96 (2): 127-156.
Some other recommended literature is cited in this paper.