(sampled with Pitfall & Colour Traps 1984-1986)
Summary and correction of the list of species by Rose (1994) - Diploma thesis, University Oldenburg, 74pp.
Zur Besiedlung der jungen Düneninsel Memmert durch Kurzflügler (Coleoptera; Staphylinidae)
[Colonization of the young dune island Memmert by staphylinid beetles (Coleoptera: Staphylinidae)]
Abstract: Between 1984 and 1986, 5874 staphylinid specimens out of 159 species were caught in pitfall traps and colour dishes on Memmert island, 17 of which not having been recorded on the old East Frisian Islands before. The most abundant species were Falagria sulcatula, Drusilla canaliculata, and Atheta fungi. Phenologies of these and some other species are shown. Ten species are typical coastal inhabitants, concerning Carpelimus foveolatus, Anotylus maritimus, Bledius spectabilis, Bl. fergussoni, Bl. subniger, Philonthus salinus, Gabrius keysianus, Quedius simplicifrons, Brundinia marina, and Atheta varendorffiana. Four main habitats were investigated of which the typical grey dunes proved to be richest in species (102 spp. in pitfall traps), followed by ruderal grey dunes (97), reeds (44) and upper salt marshes (34).
Furthermore, size structures of the staphylinidocenoses on Memmert island were examined. Small species were better represented in the reeds compared to the other investigated habitats. In addition, smaller species were stronger represented on Memmert, compared to Mellum island. This might be due to the further stage of succession and hence greater stability of Memmert, or due to a reduced comparability of the two investigations.
Mostly euryoecious species and inhabitants of marine influenced environments were successful in colonization. Information on assumed hydrophilous/xerophilous behaviour for the mainland (Koch 1988) of Gyrohypnus angustatus, Othius subuliformis, Philonthus marginatus, Gabrius keysianus, Ocypus ater, Heterothops quadripunctulus, Quedius simplicifrons, Mycetoporus splendidus, Sepedophilus nigripennis, Tachyporus hypnorum, Tachinus signatus, Aloconota gregaria, Geostiba circellaris, and Dinaraea angustula proved to be incorrect for Memmert island.
On three staphylinid species parasitic Laboulbeniales (Ascomycetes) were found. Quedius persimilis was proved to be a new host for Symplectromyces vulgaris; that fungus was known to live parasitic on other Quedius species.
Table 3: List of staphylinid species (excl. Scaphidiinae & Pselaphinae) found at Memmert island from 1984 to 1986 [PT: pitfall traps, CT: colour traps - D: typical grey dunes (young tertiary dunes), RD: ruderal grey dunes, R: reeds (esp. Phragmites) S: upper salt marsh - x: sparse in this area, 1: subrecedent, 2: recedent, 3: subdominant, 4: dominant, 5: eudominant {for dominances only catches from pitfall traps}]
| Species | PT | CT | all | D | RD | R | S |
| Megarthrus depressus (PAYK. 1789) sensu ASSING & SCHÜLKE 1999 | 1 | 1 | x | ||||
| Proteinus laevigatus HOCHH. | 2 | 2 | x | x | |||
| Micropeplus porcatus (PAYK. 1789) | 1 | 1 | x | ||||
| Omalium rivulare (PAYK. 1789) | 18 | 5 | 23 | x | 2 | x | x |
| Omalium caesum GRAV. 1806 | 3 | 1 | 4 | x | x | ||
| Omalium italicum BERNH. 1902 | 2 | 2 | x | ||||
| Anthobium atrocephalum (GYLL. 1827) | 2 | 2 | x | ||||
| Lesteva sicula ssp. heeri FAUV. 1872 | 2 | 2 | x | ||||
| Lesteva longoelytrata (GOEZE 1777) | 4 | 4 | x | 1 | |||
| Coprophilus striatulus (F. 1792) | 1 | 1 | x | ||||
| Carpelimus rivularis MOTSCH. 1860 | 1 | 1 | x | ||||
| Carpelimus corticinus (GRAV. 1806) | 13 | 3 | 16 | x | 1 | 1 | |
| Carpelimus foveolatus (SAHLB. 1832) | 6 | 5 | 11 | x | x | x | |
| Carpelimus pusillus (GRAV. 1802) | 1 | 1 | 2 | x | |||
| Carpelimus elongatulus ER. 1839 | 4 | 4 | x | x | x | x | |
| Oxytelus migrator FAUV. 1904 | 1 | 1 | |||||
| Oxytelus laqueatus (MARSH. 1802) | 1 | 1 | |||||
| Anotylus rugosus (F. 1775) | 3 | 1 | 4 | x | x | ||
| Anotylus maritimus THOMS. 1861 | 4 | 4 | |||||
| Anotylus inustus (GRAV. 1806) | 1 | 1 | x | ||||
| Anotylus tetracarinatus (BLOCK 1799) | 4 | 5 | 9 | x | x | ||
| Platystethus arenarius (FOURCR. 1785) | 1 | 1 | |||||
| Platystethus cornutus (GRAV. 1802) | 1 | 1 | x | ||||
| Bledius spectabilis ssp. frisius LOHSE 1978 | 1 | 1 | 2 | x | |||
| Bledius opacus (BLOCK 1799) | 5 | 14 | 19 | 1 | |||
| Bledius subniger SCHNEID. 1898 | 80 | 80 | |||||
| Bledius fergussoni JOY 1912 | 57 | 57 | |||||
| Stenus clavicornis (SCOP. 1763) | 5 | 5 | x | x | |||
| Stenus boops LJUNGH 1804 | 3 | 3 | 1 | ||||
| Stenus nitens STEPH. 1833 | 3 | 3 | x | x | x | ||
| Stenus brunnipes STEPH. 1833 | 3 | 3 | x | ||||
| Stenus nigritulus GYLL. 1827 | 2 | 2 | x | x | |||
| Stenus latifrons ER. 1839 | 1 | 1 | x | ||||
| Stenus fulvicornis STEPH. 1833 | 1 | 1 | x | ||||
| Stenus impressus GERM. 1824 | 1 | 1 | x | ||||
| Euaesthetus bipunctatus (LJUNGH 1804) | 5 | 5 | x | 1 | |||
| Rugilus orbiculatus (PAYK. 1789) | 2 | 9 | 11 | x | |||
| Lathrobium volgense HOCHH. 1851 | 3 | 3 | x | x | |||
| Lathrobium brunnipes (F. 1792) | 11 | 11 | x | 1 | |||
| Lathrobium impressum HEER 1841 | 4 | 4 | x | x | x | ||
| Lathrobium longulum GRAV. 1802 | 1 | 1 | x | ||||
| Cryptobium fracticorne (PAYK. 1800) | 6 | 6 | 1 | x | |||
| Cryptobium collare RTT.1884 | 1 | 1 | 2 | x | |||
| Gyrohypnus angustatus STEPH. 1833 | 45 | 2 | 47 | 2 | 1 | 1 | |
| Xantholinus linearis (OL. 1795) | 76 | 76 | 3 | 1 | x | x | |
| Xantholinus longiventris HEER 1839 | 36 | 36 | x | 1 | 1 | 2 | |
| Megalinus glabratus (GRAV. 1802) | 12 | 2 | 14 | 1 | 1 | ||
| Othius punctulatus (GOEZE 1777) | 10 | 10 | x | 1 | |||
| Othius angustus angustus STEPH. | 12 | 12 | 1 | 1 | |||
| Othius subuliformis STEPH. | 31 | 1 | 32 | 1 | 2 | ||
| Philonthus atratus (GRAV. 1802) | 4 | 4 | x | x | x | ||
| Philonthus cognatus STEPH. 1832 | 2 | 1 | 3 | x | |||
| Philonthus mannerheimi FAUV. 1869 | 4 | 2 | 6 | x | x | x | |
| Philonthus succicola THOMS. 1860 | 1 | 5 | 6 | x | |||
| Philonthus sordidus (GRAV. 1802) | 8 | 8 | |||||
| Philonthus spermophili GANGLB. 1897 | 1 | 1 | |||||
| Philonthus carbonarius (GRAV. 1810) | 10 | 1 | 11 | 1 | x | ||
| Philonthus cruentatus (GMEL. 1789) | 4 | 1 | 5 | x | x | ||
| Philonthus quisquilarius (GYLL. 1810) | 1 | 1 | x | ||||
| Philonthus salinus KIESW. 1844 | 3 | 3 | |||||
| Philonthus longicornis STEPH. 1832 | 1 | 1 | x | ||||
| Philonthus micans (GRAV. 1802) | 20 | 3 | 23 | 2 | 1 | ||
| Philonthus marginatus (STRÖM 1768) | 8 | 3 | 11 | 1 | x | ||
| Gabrius trossulus (NORDM. 1837) | 4 | 4 | x | x | |||
| Gabrius keysianus SHP. 1910 | 37 | 4 | 41 | x | 2 | 1 | 1 |
| Gabrius breviventer (SPERK) | 5 | 2 | 7 | x | x | x | |
| Gabrius subnigritulus (RTT. 1909) | 2 | 2 | x | ||||
| Ocypus brunnipes (F. 1781) | 107 | 107 | 3 | 3 | 1 | ||
| Ocypus aeneocephalus (DEGEER 1774) | 1 | 1 | x | ||||
| Ocypus ater (GRAV. 1802) | 186 | 186 | 4 | 1 | x | 1 | |
| Ocypus compressus (MARSH. 1802) | 10 | 10 | 1 | 1 | |||
| Ocypus melanarius (HEER 1839) | 3 | 3 | x | x | |||
| Heterothops quadripunctulus (GRAV. 1806) | 11 | 11 | 1 | ||||
| Quedius fuliginosus (GRAV. 1802) | 1 | 1 | 2 | x | |||
| Quedius levicollis (BRULLÉ 1832) | 23 | 23 | 1 | x | 1 | ||
| Quedius simplicifrons FAIRM. 1861 | 6 | 6 | x | 1 | |||
| Quedius molochinus (GRAV. 1806) | 46 | 46 | 2 | 1 | 1 | 1 | |
| Quedius scintillans (GRAV. 1806) | 10 | 5 | 15 | 1 | x | ||
| Quedius semiaeneus (STEPH. 1833) | 1 | 1 | x | ||||
| Quedius semiobscurus (MARSH. 1802) | 16 | 16 | 1 | x | x | ||
| Quedius persimilis MULS. REY 1875 | 31 | 31 | 2 | x | x | ||
| Mycetoporus baudueri MULS. REY 1875 | 6 | 3 | 9 | x | x | x | |
| Mycetoporus lepidus (GRAV. 1802) | 8 | 2 | 10 | x | 1 | x | |
| Mycetoporus longulus MANNH. 1830 | 1 | 1 | x | ||||
| Ischnosoma splendidum (GRAV. 1806) | 97 | 97 | 3 | 1 | 1 | 1 | |
| Bolitobius castaneus (STEPH. 1832) | 5 | 2 | 7 | x | x | ||
| Sepedophilus marshami (STEPH. 1832) | 32 | 32 | 2 | 1 | |||
| Sepedophilus nigripennis (STEPH. 1832) | 121 | 2 | 123 | 4 | 1 | x | 1 |
| Tachyporus nitidulus (F. 1781) | 14 | 1 | 15 | 1 | 1 | x | |
| Tachyporus obtusus (L. 1767) | 2 | 1 | 3 | x | |||
| Tachyporus solutus ER. 1839 | 6 | 7 | 13 | x | 1 | ||
| Tachyporus hypnorum (F. 1775) | 46 | 12 | 58 | 3 | x | ||
| Tachyporus dispar (PAYK. 1789) | 122 | 8 | 130 | 3 | 3 | 1 | x |
| Tachyporus atriceps STEPH. 1832 | 4 | 4 | 1 | ||||
| Tachyporus pulchellus MANNH. 1841 | 1 | 1 | x | ||||
| Tachyporus pusillus GRAV. 1806 | 6 | 6 | 1 | x | |||
| Lamprinodes saginatus (GRAV. 1806) | 1 | 1 | x | ||||
| Tachinus signatus GRAV. 1802 | 109 | 4 | 113 | x | 4 | ||
| Tachinus laticollis GRAV. 1802 | 5 | 4 | 9 | 1 | |||
| Tachinus marginellus (F. 1781) | 6 | 8 | 14 | 1 | |||
| Tachinus corticinus GRAV. 1802 | 110 | 110 | 1 | 4 | 3 | ||
| Myllaena intermedia ER. 1837 | 2 | 2 | x | ||||
| Myllaena minuta (GRAV. 1806) | 7 | 2 | 9 | 1 | |||
| Myllaena infuscata KR. 1853 | 1 | 1 | x | ||||
| Oligota pusillima (GRAV. 1806) | 25 | 25 | 2 | x | 1 | ||
| Cypha tarsalis (LUZE 1902) | 3 | 3 | x | x | |||
| Autalia rivularis (GRAV. 1802) | 2 | 2 | |||||
| Falagria sulcatula (GRAV. 1806) | 2279 | 1 | 2280 | 1 | 5 | 5 | 5 |
| Tachyusa constricta (ER. 1837) | 1 | 1 | x | ||||
| Aloconota gregaria (ER. 1839) | 50 | 12 | 62 | 2 | 3 | x | |
| Amischa analis (GRAV. 1802) | 38 | 2 | 40 | 2 | 1 | x | x |
| Amischa nigrofusca (STEPH. 1832) | 1 | 4 | 5 | x | |||
| Amischa decipiens (SHP. 1869) | 1 | 1 | x | ||||
| Brundinia marina (MULS. REY 1853) | 2 | 2 | 4 | x | x | ||
| Ousipalia caesula (ER. 1839) | 18 | 18 | 1 | ||||
| Geostiba circellaris (GRAV. 1806) | 44 | 44 | 1 | 3 | x | 1 | |
| Dinaraea angustula (GYLL. 1810) | 7 | 7 | x | 1 | x | ||
| Atheta elongatula (GRAV. 1802) | 5 | 2 | 7 | x | x | ||
| Atheta palustris (KIESW. 1844) | 1 | 1 | x | ||||
| Atheta divisa (MÄRK. 1844) | 3 | 9 | 12 | x | |||
| Atheta amicula (STEPH. 1832) | 1 | 1 | |||||
| Atheta orbata (ER. 1837) | 12 | 6 | 18 | 1 | |||
| Atheta fungi (GRAV. 1806) | 267 | 11 | 278 | 5 | 4 | 1 | x |
| Atheta amplicollis (MULS. REY 1873) | 1 | 1 | 1 | x | |||
| Atheta celata (ER. 1837) | 2 | 2 | |||||
| Atheta triangulum (KR. 1856) | 10 | 3 | 13 | 1 | x | x | |
| Atheta xanthopus (THOMS. 1856) | 1 | 1 | |||||
| Atheta aeneicollis (SHARP 1869) | 3 | 1 | 4 | x | x | ||
| Atheta repanda (MULS. REY 1874) | 1 | 1 | 2 | x | |||
| Atheta macrocera (THOMS. 1856) | 2 | 2 | x | ||||
| Atheta nigripes (THOMS. 1856) | 1 | 1 | 2 | x | |||
| Atheta varendorffiana BRNH. SCHRP. 1926 | 1 | 1 | |||||
| Acrotona exigua (ER. 1837) | 21 | 21 | x | 2 | |||
| Acrotona pygmaea (GRAV. 1802) | 7 | 7 | 1 | 1 | |||
| Acrotona aterrima (GRAV. 1802) | 1 | 2 | 3 | x | |||
| Drusilla canaliculata (F. 1787) | 803 | 803 | 5 | 5 | 3 | 5 | |
| Zyras limbatus (PAYK. 1789) | 1 | 1 | x | ||||
| Ilyobates nigricollis (PAYK. 1800) | 1 | 1 | x | ||||
| Calodera aethiops (GRAV. 1802) | 5 | 5 | |||||
| Calodera protensa MANNH. 1830 | 1 | 1 | x | ||||
| Meotica exilis (ER. 1837) | 2 | 1 | 3 | x | |||
| Meotica exillima SHP. 1915 | 1 | 1 | |||||
| Meotica capitalis (MULS. REY 1873) | 1 | 3 | 4 | x | |||
| Oxypoda procerula MANNH. 1830 | 10 | 10 | 1 | x | x | ||
| Oxypoda opaca (GRAV. 1802) | 2 | 2 | x | x | |||
| Oxypoda longipes MULS. REY 1861 | 5 | 5 | 1 | ||||
| Oxypoda acuminata (STEPH. 1832) | 1 | 1 | x | ||||
| Oxypoda lentula ER. 1837 | 1 | 1 | x | ||||
| Oxypoda brevicornis (STEPH. 1832) | 1 | 1 | x | ||||
| Oxypoda exoleta ER. 1839 | 15 | 15 | 1 | x | |||
| Oxypoda praecox ER. 1839 | 185 | 185 | 4 | 3 | 1 | 2 | |
| Oxypoda brachyptera (STEPH. 1832) | 19 | 19 | 1 | 1 | x | ||
| Oxypoda haemorrhoa (MANNH. 1830) | 4 | 4 | x | x | |||
| Tinotus morion (GRAV. 1802) | 2 | 1 | 3 | x | |||
| Aleochara curtula (GOEZE 1777) | 3 | 9 | 12 | x | x | ||
| Aleochara lanuginosa GRAV. 1802 | 2 | 2 | x | x | |||
| Aleochara cuniculorum KR. 1858 | 11 | 5 | 16 | 1 | x | ||
| Aleochara bipustulata (L. 1761) | 2 | 2 | x | ||||
| Polystomota punctatella (MOTSCH. 1858) | 1 | 1 | |||||
| Specimens | 5493 | 381 | 5874 | 1834 | 1235 | 1308 | 1106 |
| Species | 142 | 74 | 159 | 102 | 97 | 34 | 44 |